Derek the Hybrid Detective

DD0217 DNA / New genera

Ed: "Recent DNA analysis has shown many errors in the way Bromeliads have been grouped together in genera.
It has mostly been Tillandsia studied so-far.
In the past a lot of emphasis was placed on flower structure to group into genera, but it has long been suspected this was locking us into mistakes.
Rather than simply transferring species among the old genera, new genera names are being adopted for a clean start.

Interestingly some of my favourites, the intergenerics, will no longer be intergenerics and some plain crosses will become intergenerics !

It will take many years for the new groupings to be adopted.
Past naming corrections are still not adopted after many decades in some collections !
To keep references precise, it is always wise to check for past, alternate, and corrected names of any new plants you get by checking on http://bromeliad.nl/taxonlist/, the 'Taxon List', and following the name through all it's changes.
Some species have been swapped around between Tillandsia, Vriesea, etc, some have new genera groups.
Here is a table of some of the latest Tillandsioideae DNA re-groupings, as supplied by Derek Butcher 01/02/17, from Barfuss."

Note: see end of this page for generic definition of the new genus Goudaea, Bt W.Till & Barfuss.


There are 8 tables below.
The first two are new names for Tillandsia/Vriesea species, then consequential name changes for 'T/V Cultivars'. (Feb. 2017)
These are followed by Orthophytum/Sincoraea species, then consequential name changes for the 'Cultivars'. (March 2017)
These are followed by Cryptanthus species (Sept'2017). The resultant 'Cultivar/Bigeneric' changes not yet available.
The new Cipuropsis species
Proposed raising of Tillandsia sub-genus Viridantha to genus level.

SpeciesOld GenusNew Genus
acostasolisii
Tillandsia
Lemeltonia
amadoi
Vriesea
Stigmatodon
anceps
Tillandsia
Wallisia
apparicianus
Vriesea
Stigmatodon
asplundii
Tillandsia
Josemania
belloi
Vriesea
Stigmatodon
bibeatricis
Vriesea
Lutheria
bifidus
Vriesea
Stigmatodon
brassicoides
Vriesea
Stigmatodon
brownie
Mezobromelia
Gregbrownia
chrysostachys
Vriesea
Goudaea
chrysostachys var stenophylla
Vriesea
Goudaea
cornuta
Tillandsia
Lemeltonia
costae
Vriesea
Stigmatodon
croceanus
Vriesea
Stigmatodon
cyanea
Tillandsia
Wallisia
delicatula
Tillandsia
Josemania
dodsonii
Tillandsia
Lemeltonia
dyeriana
Tillandsia
Racinaea
euclidianus
Vriesea
Stigmatodon
fontellanus
Vriesea
Stigmatodon
fulgens
Mezobromelia
Gregbrownia
funebris
Vriesea
Stigmatodon
gastinianus
Vriesea
Stigmatodon
glutinosa
Vriesea
Lutheria
goniorachis
Vriesea
Stigmatodon
grandis
Tillandsia
Pseudalcantarea
hamaleana
Tillandsia
Racinaea
harrylutheri
Vriesea
Stigmatodon
heliconioides
Vriesea
Tillandsia
hutchisonii
Mezobromelia
Gregbrownia
laxissima
Tillandsia
Barfussia
laxissima var moorei
Tillandsia
Barfussia
lindeniana (new, was umbellata)
Tillandsia
Wallisia
lymansmithii
Mezobromelia
Gregbrownia
macropetala
Tillandsia
Pseudalcantarea
magnibracteatus
Vriesea
Stigmatodon
malzinei
Vriesea
Tillandsia
monadelpha
Tillandsia/
Lemeltonia
monstrum
Vriesea
Jagrantia
multifoliatus
Vriesea
Stigmatodon
narthecioides
Tillandsia
Lemeltonia
ospinae
Vriesea
Goudaea
ospinae var gruberi
Vriesea
Goudaea
pinnata
Tillandsia
Josemania
platyrhachis
Tillandsia
Barfussia
plurifolius
Vriesea
Stigmatodon
pretiosa
Tillandsia
Wallisia
rosulatulus
Vriesea
Stigmatodon
sanctateresensis
Vriesea
Stigmatodon
scaligera
Tillandsia
Lemeltonia
singularis
Tillandsia
Josemania
soderstromii
Vriesea
Lutheria
splendens
Vriesea
Lutheria
splendens var chlorostachya
Vriesea
Lutheria
splendens var formosa
Vriesea
Lutheria
splendens var oinochroma
Vriesea
Lutheria
splendens var striatifolia
Vriesea
Lutheria
triglochinoides
Tillandsia
Lemeltonia
truncate
Tillandsia
Josemania
truncata var major
Tillandsia
Josemania
tuerckheimii
Vriesea
Zizkaea
venusta
Tillandsia
Racinaea
viridiflora
Tillandsia
Pseudalcantarea
wagneriana
Tillandsia
Barfussia
x duvalii treat as ‘Duvalii’
Tillandsia
Wallisia

BCR Cultivars: genus changes generated via Tillandsioideae DNA Studies, Barfuss et al Paper 2016. Published in Phytotaxa 279 (1), October, 2016.
Cultivars compiled by Geoff Lawn, BSI Cultivar Registrar, March 2017

CultivarNEW GenusOLD Genus
Batik
Goudaea
Vriesea
Espirito Santo
Goudaea
Vriesea
Euro Smudge
Goudaea
Vriesea
Fortuna
Goudaea
Vriesea
Golden Zombie
Goudaea
Vriesea
Hawaiian Magic
Goudaea
Vriesea
Hot Chocolate
Goudaea
Vriesea
Jersey Caramel
Goudaea
Vriesea
Josefa Leon
Goudaea
Vriesea
Nazca
Goudaea
Vriesea
Santa Maria
Goudaea
Vriesea
Scaredy Cat
Goudaea
Vriesea
Smudge Bob
Goudaea
Vriesea
Smudge Grub
Goudaea
Vriesea
Sons of Tiger Tim
Goudaea
Vriesea
Tiger Tim
Goudaea
Vriesea
White Zombie
Goudaea
Vriesea
Witchetty Grub
Goudaea
Vriesea
Dreamchaser
x Guzgoudaea
x Guzvriesea
Jags Hunua Fire
x Vriesgoudaea
Vriesea
Jags Hunua Fountain
x Vriesgoudaea
Vriesea
Jags Hunua Gloss
x Vriesgoudaea
Vriesea
LSD
x Vriesgoudaea
Vriesea
Pactole
x Vriesgoudaea
Vriesea
Scarlet Fountain
x Vriesgoudaea
Vriesea
Stolen Glory
x Vriesgoudaea
Vriesea
Andreana
x Vrieslutheria
Vriesea
Deroose
x Vrieslutheria
Vriesea
Dragon
x Vrieslutheria
Vriesea
Elmireana
x Vrieslutheria
Vriesea
Enid Frase
x Vrieslutheria
Vriesea
Henrici
x Vrieslutheria
Vriesea
Insignis H.L.B.
x Vrieslutheria
Vriesea
Kent’s Sunset
x Vrieslutheria
Vriesea
Louis Dutrie
x Vrieslutheria
Vriesea
Pommero Escheana
x Vrieslutheria
Vriesea
Psittacino-Splendens
x Vrieslutheria
Vriesea
Sphinx
x Vrieslutheria
Vriesea
Symphonie
x Vrieslutheria
Vriesea
Towering Flame
x Vrieslutheria
Vriesea
Ubanteniana
x Vrieslutheria
Vriesea
Wanteniana
x Vrieslutheria
Vriesea
Witteana
x Vrieslutheria
Vriesea
Pink Magic
x Barvriesea
x Vrieslandsia
Magnifica
x Guzlutheria
x Guzvriesea
Mirabilis
x Guzlutheria
x Guzvriesea
Splendulata
x Guzlutheria
x Guzvriesea
Carlsbad
x Luthandsia
Vriesea
Leopoldiana
x Luthandsia
Vriesea
Brasilia
Lutheria
Vriesea
Cayenne
Lutheria
Vriesea
Chantrieri
Lutheria
Vriesea
Double Pleasure
Lutheria
Vriesea
Double Pleasure Harlequin
Lutheria
Vriesea
Fascination
Lutheria
Vriesea
Favoriet
Lutheria
Vriesea
Fire
Lutheria
Vriesea
Flaming Sword
Lutheria
Vriesea
Flammendes Schwert
Lutheria
Vriesea
Galaxy
Lutheria
Vriesea
Formosa
Lutheria
Vriesea
Hera
Lutheria
Vriesea
Illustris
Lutheria
Vriesea
Johnson’s #637
Lutheria
Vriesea
Juno
Lutheria
Vriesea
Madam Pele
Lutheria
Vriesea
Major
Lutheria
Vriesea
Meyer’s Favorite
Lutheria
Vriesea
Mortfontanensis Splendens
Lutheria
Vriesea
Nigre
Lutheria
Vriesea
Splendide
Lutheria
Vriesea
Splendide Vista
Lutheria
Vriesea
Splenriet
Lutheria
Vriesea
Tepuy
Lutheria
Vriesea
Trinidad
Lutheria
Vriesea
Silver Candelabra
Pseudalcantarea
Tillandsia
La Mano Magica
Racinaea
x Racindsia
Kiseki
x Racindsia
Tillandsia
Miracle
x Racindsia
Tillandsia
Mirai
x Racindsia
Tillandsia
Sexy Pink
Barfussia
Tillandsia
Mira
Tillandsia
Vriesea
Amanda Isley
x Vrieslandsia
Vriesea
Fernanda
x Vrieslandsia
Vriesea
Groves Big Red
x Vrieslandsia
Vriesea
Kent’s Surprise
x Vrieslandsia
Vriesea
Talon
x Vrieslandsia
Vriesea
Anita
Wallisia
Tillandsia
Angela
Wallisia
Tillandsia
Anwyl Expo
Wallisia
Tillandsia
Ariane
Wallisia
Tillandsia
Caeca
Wallisia
Tillandsia
Carmen
Wallisia
Tillandsia
Duvaliana
Wallisia
Tillandsia
Duvalii
Wallisia
Tillandsia
Emilie
Wallisia
Tillandsia
Hans Gulz
Wallisia
Tillandsia
Hybride H G
Wallisia
Tillandsia
Josee
Wallisia
Tillandsia
Leo
Wallisia
Tillandsia
Nicoline
Wallisia
Tillandsia
Paradise
Wallisia
Tillandsia
Pink Plume
Wallisia
Tillandsia
Roku
Wallisia
Tillandsia
Sandy
Wallisia
Tillandsia
Triflor
Wallisia
Tillandsia
Holm’s Deep Purple
x Wallandsia
Tillandsia
Creation
x Wallfussia
Tillandsia
Feather Belle
x Wallfussia
Tillandsia
Hana
x Wallfussia
Tillandsia
Hercules
x Wallfussia
Tillandsia
Lynnie
x Wallfussia
Tillandsia
Tropic Delight
x Wallfussia
Tillandsia
Tropic Torch
x Wallfussia
Tillandsia
Babylon
x Zizkagoudaea
V. tuerckheimii x Tiger Tim



Genera Changes: Species: Orthophytum / Sincoraea March 2017.
Reference: BSI Journal Vol.66(1) pp.6-19 (Jan-March, 2016). Re-establishment of Sincoraea (Bromeliaceae). R Louzada & M Wanderley.

From Geoff Lawn ... "Only these 11 Orthophytum species are reclassified as Sincoraea species--all are "sunburst" types with sessile inflorescences (no stem or peduncle), in contrast to the "pendunculate" (visible flower stem.) of other species in the genus Orthophytum.
A key to Sincoraea species is presented in the BSI Journal, botanical descriptions of all 11 species and their synonyms. Because a number of sunburst types have been discovered and described in the last 20 years or more, botanists have gotten a better idea of how best to classify the various clades of 57 species of Orthophytum + 11 Sincoraea species.

Herb Plever has written a good summary of the logic behind these changes in the March 2017 issue of 'BROMELIANA' published by the New York Bromeliad Society.
Click this LINK for a copy of the article, including identification Key.

speciesNEW GenusOLD Genus
albopicta
Sincoraea
Orthophytum
amoena
Sincoraea
Orthophytum
burle-marxii
Sincoraea
Orthophytum
hatschbachii
Sincoraea
Orthophytum
heleniceae
Sincoraea
Orthophytum
humilis
Sincoraea
Orthophytum
mucugensis
Sincoraea
Orthophytum
navioides
Sincoraea
Orthophytum
ophiuroides
Sincoraea
Orthophytum
rafaelii
Sincoraea
Orthophytum
ulei
Sincoraea
Orthophytum

Genera Changes: Cultivars & new bigenerics: Orthophytum / Sincoraea. Compiled by Geoff Lawn, BSI Cultivar Registrar, March 2017.

Andromeda
x Sincoregelia
x Neophytum
Aurora
x Sincoregelia
x Neophytum
Blushing Bride
x Sincoregelia
x Neophytum
Burgundy Hill
x Sincoregelia
x Neophytum
Burgundy Thrill
x Sincoregelia
x Neophytum
Cosmic Blast
x Sincoregelia
x Neophytum
Ecstasy
x Sincoregelia
x Neophytum
Firecracker
x Sincoregelia
x Neophytum
Galactic Warrior
x Sincoregelia
x Neophytum
Gary Hendrix
x Sincoregelia
x Neophytum
George H Anderson
x Sincoregelia
x Neophytum
Lisanne Kiehl
x Sincoregelia
x Neophytum
Lymanii
x Sincoregelia
x Neophytum
Medalist
x Sincoregelia
x Neophytum
Mollie S
x Sincoregelia
x Neophytum
Ralph Davis
x Sincoregelia
x Neophytum
Rising Tide
x Sincoregelia
x Neophytum
Shiraz
x Sincoregelia
x Neophytum
Supernova
x Sincoregelia
x Neophytum
Andrea
Sincoraea
Orthophytum
Blaze
x Sincorphytum
Orthophytum
Blazing Bonsai
x Sincortanthus
x Orthotanthus
Rosita
x Sincorglaziovia
x Orthoglaziovia
Selby
x Nidusincoraea
x Ortholarium
Powderpuff
x Sincoraechmea
x Orthomea



Goudaea W. Till & Barfuss, gen. nov. Phytotaxa 279(1): 001-097. 2016.
Diagnosis: A genere Vriesea Lindl. stigmate simplico-erecto, ovulis obtusis et petalis cucullatis differt.
Type: Vriesea chrysostachys E. Morren = Goudaea chrysostachys (E. Morren) W. Till & Barfuss
Description:
Plants epiphytic or terrestrial, usually medium sized, impounding rosette herbs.
Leaves mesomorphic; leaf blades lingulate, appearing glabrous.
Inflorescence simple or once branched;
floral bracts ecarinate, concealing the sepals completely; flowers distichously arranged,
Sepals symmetric; petals yellow or whitish, short connate at the base, tips cucullate, forming a tubular, hardly opened, actinomorphic corolla, bearing spathulate basal appendages, the appendage tips undivided and obtuse or incised and acute;
stamens shorter than the petals, included within the corolla;
filaments short adnate to the connate portion of the petals;
anthers subbasifixed;
pollen sulcate, exine reticulate, with a sulcus of the complex, diffuse type (subtype c);
ovules obtuse; style included within the corolla; stigma of the simple-erect type, papillose.
Distribution: Columbia to central Peru.
Etymology: Named in honor of Eric John Gouda (1957-) from Utrecht, The Netherlands, long-term researcher in Bromeliaceae.
Note: Two species and two varieties.



Cryptanthoid changes due to Leme et al in Phytotaxa 318(1): 001-088. 2017, by D.Butcher Aug 2017.
There are 3 new genera – Forzzaea, Hoplocryptanthus and Rokautskyia.

Note 1. Lapanthus vidaliorum has also been moved to Hoplocryptanthus
Note 2. this oportunity has been taken to correct the latin spelling and grammar of some species.

Cryptanthus SpeciesReferenceNew Genus
aracruzensis
Leme & Kollmann
Rokautskyia
caracensis
Leme & Gross
Hoplocryptanthus
caulescens
I Ramirez
Rokautskyia
exaltatus
Luther
Rokautskyia exaltata
fernseeoides
Leme
Rokautskyia
ferrarius
Leme & Paula
Hoplocryptanthus
glaziovii
Mez
Hoplocryptanthus
latifolius
Leme
Rokautskyia latifolia
lavrasensis
Leme
Hoplocryptanthus
leopoldo-horstii
Rauh
Forzzaea
leuzingerae
Leme
Rokautskyia
microglazioui
I Ramirez
Rokautskyia
micrus
Lousada, Wand. & Versieux
Forzzaea micra
odoratissimus
Leme
Rokautskyia odoratissima
pseudoglaziovii
Leme
Rokautskyia pseudoglazioui
pseudoscaposus
L B Sm.
Rokautskyia pseudoscaposa
regius
Leme
Hoplocryptanthus
roberto-kautskyi
Leme
Rokautskyia
sanctaluciae
Leme & Kollmann
Rokautskyia
scaposus
E Pereira
Rokautskyia scaposa
schwackeanus
Mez
Hoplocryptanthus
tiradentesensis
Leme
Hoplocryptanthus
warasii
Pereira
Forzzaea
whitmanii
Leme
Rokautskyia
Lapanthus vidaliorum
(OBC Ribeiro & Paula) Louzada & Wand
Hoplocryptanthus


Cipuropsis species, as recorded in BSI Journal April-June 2016, E.J.Gouda article.

Cipuropsis SpeciesReferenceOld Genus
amicorum
(I.Ramírez & Bevilacqua) Gouda
Was Tillandsia
asmussii
Gouda
New species
subandina
Ule
Was Tillandsia, then Vriesea


Proposed Viridantha genus.
Key to identify species in sub-genus Viridantha 2018. Translated by Derek Butcher.
Derek Butcher ... "The latest to come out of Mexico is that they still persist in treating them as Viridantha at genus level and not sub-genus level. However they have produced a very comprehensive paper on the subject (In Spanish!). It is an interesting group and several are being grown in Australia. The species are not easy to recognise but the new key gives a different approach to identification. You may notice that T. yagulensis has been promoted to species level from being a variety of T. atroviridipetala. You may also notice the new T. rzedowskiana but I haven't a clue as to its pronunciation !"

1.Rosettes irregular in general outline or sickle shaped => 2
1.Rosettes spherical in general outline => 6

2.Inflorescence compound => Viridantha boqueronensis
2.Inflorescence simple => 3

3.Rosettes sickle shaped, leaves curved => 4
3.Rosettes irregular, leaves scaled => 5

4.Plants usually greater than 5.1 cm high; peduncle more than 2.1 cm long => Viridantha curvifolia
4. Plants usually less than 5 cm high; peduncle less than 2.1 cm long. => Viridantha penascoensis

5. Peduncle more than 4.l cm long and less than 2mm diam => Viridantha tortilis
5. Peduncle less than 4 cm ong and more than 4 mm diam => Viridantha lepidosepala

6. Peduncle more than 4.1 cm long. => 7
6.Peduncle less than 4 cm long. => 9

7. Inflorescence with a single spike, more than 3.1 cm long, applanate to cylindrical, rarely with 2 shorter lateral spikes and adpressed to the principal => Viridantha ignesiae
7. Inflorescence with 2 or more spikes, less than 3 cm long applanate => 8

8. Width of leaf at base larger than 2.1 mm; floral bracts triangular to ovate, carinate for entire length abaxially => Viridantha plumosa
8. Width of leaf at base less than 2 mm; floral bracts elliptic-oblong, carinate only from centre to tip abaxially => Viridantha caballosensis

9. Leaf blade more than 7.1 mm wide; anthers generally black => Viridantha mauryana
9. Leaf blade less than 7 mm wide; anthers green to browner => 10

10. Plant short caulescent; floral bracts elliptic => Viridantha yagulensis
10. Plant acaulis; floral bracts ovate to amply ovate, lanceolate or triangular => 11

11. Spike more than 3.1 cm long and more than 1 cm wide => 12
11. Spike less than 3 cm long and less than 1 cm wide => 13

12. Leaf sheath amply ovate to oblong, blade more than 4 mm wide, sepals less than 1.6 cm => Viridantha rzedowskiana
12. Leaf sheath closely oblong, blade less than 4 mm wide; larger of the sepals to 1.5 cm => Viridantha grandispica

13. Leaf blade more than 4 mm wide; spikes generally less than 3; floral bracts less than 1.5 cm long => Viridantha teloloapanensis
13. Leaf blade less than 4 mm wide; spikes generally more than 3; floral bracts more than 1.5 cm long => Viridantha atroviridipetala



New genus Karawata by Derek Butcher Sept 2019
This new genus was created by J. Marciel et al. in Systematic Botany 44(3): 519–535. 2019
Phylogenetics work revealed that Aechmea subg. Chevaliera is not a monophyletic group. Seven species previously assigned to the subgenus form a clade with strong statistical support and in sister position to morphologically distinct members of other genera. Morphological and phylogenetic evidence segregates these seven species in a new genus named Karawata, which requires the following new combinations: Karawata depressa, Karawata gustavoi, Karawata hostilis, Karawata multiflora, Karawata nigribracteata, Karawata prasinata, and Karawata saxicola.

These days the trend seems to be to create new genera rather than trying to solve the problem at sub-genus level. This in turn creates problems with naming of man-made hybrids. In this case a quick check of the Bromeliad Cultivar Register has revealed that none of the 7 taxa have been reported. All named have impressive inflorescences but are large plants and take many years to flower which may be the cause of reluctance to hybridise.

Karawata multiflora by Oscar Ribeiro

Updated 03/09/19